>>>The proof marks I see are from the period 1891-1939.The DDR adopted very similar marks, causing your confusion.<<<
No sir, when you compare the proof marks from the period 1891 - 1939 with the DDR marks than you can see the crown and the letters are different in form.
I have compared the shown marks with the proof marks on my Nimrod side by side shotgun, manufactured 1921 in Suhl and I'm absolutely shure this gun in question was manufactured in DDR after WWII by Ernst Thälmann Werke, formerly Fortuna Werke and formerly Sauer & Sohn. This factory was manufacturing Sauer & Sohn armory until end of 1960th and Sauer & Sohn, Eckernförde, Western Germany stand in action against and win at least.
I think the mark "MADE IN PRUSSIA" was made by the importer, to get a higher price, never seen in Germany.